Key botanical
characters:
Evergreen
dioecious trees, to 15 m high, bark black or grey-black, rough, peeling off
in small rectangular pieces, fissured, brittle; blaze yellow; branchlets
slender, apical part sparsely minutely adpressedpilose or glabrous. Leaves
simple, alternate, estipulate; petiole 4-10 mm long, slender, glabrous;
lamina 4.5-15 x 4-7 cm, elliptic-oblong, oblong, oblong-lanceolate, obovate
or obovate-oblong, base cuneate or acute, apex obtuse, emarginate or
subacute, margin entire, glabrous, coriaceous; lateral nerves 5-11 pairs,
pinnate, prominent, basal pairs opposite, intercostae reticulate, prominent,
pellucid. Flowers unisexual, subsessile, greenish-yellow; male flowers: 3-15
in short, axillary, umbellate clusters; peduncle densely tomentose; pedicel 3
mm; calyx cupular, glabrous; lobes 4, ovate, spreading, margin ciliate;
corolla 7 mm long, 1 cm across, tubular to salver-shaped; lobes 4,
ovate-oblong, 6 mm long; stamens 6-12, unequal, in uneven groups; filaments
1.5-3.5 mm long; anthers linear, connectives crested, apiculate; pistillode
linear to 2 mm; female flowers: solitary, axillary, pedicel to 3 mm; calyx
cupular, mouth densely tomentose; lobes 4, ovate, spreading, subacute;
corolla 3 mm across, tubular; tube 6 mm; lobes 4, each 6 mm, acute, reflexed;
staminodes 8 with double anthers; ovary superior, 4.5 x 4 mm, globose,
8-celled, ovules 1-in each cell; styles 4; stigma capitellate. Fruit a berry,
2 x 1.8 cm, subglobose, with short apical beak; calyx enlarged and forming a
shallow wooden cup; reflexed; seeds 3-8, black; endosperm equable.
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